http:/www.paper.edu.cn 1,2 , 2 , 2 , 1 ( 1 , 400716; 2 , 100094) E-mail lidongchaolihaoyahoo.com.cn S499 1-3 1 4-7 Gause 4 8-10 6, 11 Gause 12 B A ZHJ-1 B 13, 14 15, 16 1978 DeBach “ 17 ” DeBach “973” 2002CB111400 -1- http:/www.paper.edu.cn 18 2002 Reitz DeBach 19 19, 20 19 Pineus coloradensis Pineus boerneri P . boerneri P . coloradensis P . boerneri -2- http:/www.paper.edu.cn 35 Fiorina externa Nuculaspis (Tsugaspidotus) tsugae F . ex