作物学报 2017, 43(3): 332342 ; SE1S (31471491, 31501310), SEy3S/v (2014Sj , , , * , , , 15= . lxh Hh=hYF . T , 2014, 40: 15571564 D, M, H, , , F, C. of to in 2014, 40: 15571564 (13 , M. to in 2009, 118: 631642 14 , , v* , o , fo , 9R , _, , ): , m . lxhFiiy T , 1995, 21: 257262 L, D, J, , Z, Q, J, Y, L. a to 1995, 21: 257262 (15 G, Y, J, Y, , L. in a to 2009, 118: 10591064 16 L, J, J, Y, X, J, X. of 2009, 118: 11731180 17 X, R, J, , J, P, Q, Y, K, J, J. of in a 2014, 9: 18 Q, L, T, J, J, M, , , X, Y. of a on BL 2009, 119: 531539 19 , J, , J, M, L, Y, X, Y. of a 2009, 119: 223230 20 Q, G, H, , X, Y, Q. of a on D of 2011, 123: 10991106 21 G, J, F, M, X, J. of on BS to in 2013, 126: 13971403 22 , Y, , Y, L, J, J, Q. of a in 2012, 125: 14251432 23 T, X, F, H, M, L, T, Q, L, G. of a 42 T 43 2015, 128: 613622 24 X, J, T, M, Y, F, T, G. of a 2015, 128: 20772084 25 , . 8luk1js . j . Yljv 399 Fxhy s0S: . L. , 2013, 14: 104108 , G, B, , , Z, H, X, M, X, Y. in J 2013, 14: 104108 (29 , , * , u, , k , lc , . lj 14 Fxhy8 . T . 2014, 40: 798804 , G, L, H, M, F, J. of to in 4. 2014, 40: 798804 (30 G, , L, M, J, W, M, F, H, J. to in 55: of 2015, 55: 10171025 31 T, X, X, F, W, T, G. in m2 of 00. 2015, 35: 110 32